Spanish rider Juan Ayuso pulled on the leader's yellow jersey on Tuesday after stage three of the Paris-Nice cycling race.

INEOS Grenadiers, led by Kevin Vauquelin and Oscar Onley, won the team time trial in Pouilly-sur-Loire with Ayuso's Lidl-Trek team finishing second.

Frenchman Vauquelin powered the British squad to victory in the 23.5km test through the Nièvre region of central France, giving INEOS a two-second advantage over Lidl-Trek.

Decathlon AG2R La Mondiale finished third at 11 seconds while Jonas Vingegaard's Team Visma-Lease a Bike placed fourth at 15 seconds.

Juan Ayuso Leads Tight General Classification Battle

Ayuso took the yellow jersey from American Luke Lamperti thanks to four bonus seconds he gained in Monday's intermediate sprint.

The Spaniard ended the day two seconds ahead of Vauquelin and three seconds clear of Onley, who climbed from 63rd overall after the team effort.

Stage four will be a 195km route from Bourges to Uchon with an uphill finale marking the first summit finish of this year's "Race to the Sun," which concludes Sunday in Nice.